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Anyway, they open the package and inside the device now in proximity to its intended location activates. It turns out that it was a detonator for the nukes, it sets off the nukes from inside the silos, cracking the earth from the inside out. The tunnelers living deep below the surface from fev and radiation mutation now have access to the surface and infest the place.
Ulysses claims that the courier in new Vegas was the courier who delivered the package. He wants revenge because he's crazy, but what he really wants is you to witness him nuke the ncr so the legion can take over and collapse in itself as is inevitable, and resets fallout back to post apocalypse. (the main writer for this specific dlc and character has gone on record saying he wanted to renuke the fallout universe, but the lead director of new Vegas and probably Bethesda as well didn't share this view. This dlc was pretty much him getting his little say, and kind of projecting what he wants in the franchise as result of Ulysses actions if he's successful).
This is left open enough in the game to be either the courier we play as, or someone else who Ulysses has mistaken you with, or even Ulysses himself who can't admit his failure and projects it onto someone else.
Though if you think about it, the underlying theme of this dlc is "don't shoot the messenger".
The NCR came later, when they were searching for a route to expand; since the I-15 was the only way out of California that was not a gigantic northern detour, they had to go through there. This is when someone (Ulysses implies Caesar) sent an old piece of Remnants tech, a salvaged detonator for the pre-war nukes, to be delivered to Hopeville. It was a proximity detonator, so when the Courier arrived in the area, all nukes with active receivers went off. This also wrecks large parts of the road from California to Nevada.
As for Ulysses, he was supposed to be spying on the place. He liked the settlement and the society that went there, reminded him of his old tribe. Then a small package destroyed all of it, and it made him think in new ways of the world around him.
If you play the parts where he thinks you are NCR, it is a bit more difficult to piece together the backstory. You need to get the neutral one for the most sensible explanations.