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There have been various bugs over the years though that lead to this issue so if you are 100% sure your base is air tight, ventilators placed, oxygen tanks full and oxygen is on, then you might want to look at making a bug report.
I'm going to assume you have power because you must have Grow Lights. Is the power ever going out at any time, though? Like you run out of fuel or energy in your Capacitors.
I remember having rad issues years ago with my plants when fog rolled in but the base was airtight. It was some kind of glitch, and had to move my garden to a different spot in the base.
The other thing to look for are things like fuel tanks, generators or other blocks that give off heat or radiation less than a full block (in a cube around it) from the growplot or plant. Walk around your garden and watch for spikes in radiation or temperature. Under a grow light it should be a constant 25 C and 0 rads if air tight.