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Spectrals cluster
Only for the Spectrals civic. You start in an own cluster outside of the galaxy on a special ringworld with unique districts. Apart from that 2 other ruined ringworlds are nearby to repair.
The perk "Interstellar Overclocking" offers a unique crisis path with new structures, ships, techs and more.
Certast cluster
Only for the Certast civic.You start in an own cluster outside of the galaxy on a special ringworld with unique districts. You will find another ringworld to repair.
Requiem
Only for the Certus civic. You start with 4 special planets in your homesystem which have strong bonuses and characteristics.
Halo
You start on a giant habitat with the shape of a ring. Check the situation log, there will be a project to finish to unlock the "Halo"-megastructure. This structure is like a superweapon, you can destroy enemy planets and fleets with it!
The ascension perk "Array of the end" will unlock another Halo structure, which will be an own crisis path to win the game. (it will end in a total war and you can blow up the galaxy)
Phoenix Empire
A system with the unique planet classes "Stahl". There is also a damaged Stratetic Coordination Center in their homesystem.
Prism World
Only for Lithoids, you start on a wonderful planet made out of crystals. You will be able to increase the planet size up to 35, which will unlock a powerful deposit bonus. Use the planetary decision to let the planet "grow" larger!
Vi
For the "Vero" civic, you start in a beautifil system with the "Immortal"-class planets, which have strong bonuses for your empire. You can terraform other planets to the Immortal class right after the game begins with the help of a new ship, but it isnt so cheap to get at start.
Hiveworld
Only for hiveminds: you start on a Hiveworld.
Solarflare
This empire just had its first contact with another space empire, but the friendly neighbours just got wiped out by a solar flare! You are left with a devastated homeworld and the 3 known Fallen Empire technologies (darkmatter shields, reactor, power core) and the ability to mine dark matter deposits. One of such deposits will spawn just one jump away! Not for machines.
Ark of the constructors
Once a paradise for their organic creators, this world is now a giant structure of metal and constructions which even exceed the horizon/atmosphere. This origin is only for machines, and those machines will be able at some point to extend and use all the incredible large structures & districts there. Use the planetary decision to increase the amount of districts up to 80.
Planet size 40 will unlock a special kind of research district, which produces research points without any pops needed.
Intergalactic Isolationists (2x)
This empire lives in a star cluster outside of the galaxy. Your cluster will be connected to the galaxy via "Interstellar Portals" (all empires can use them). They will be visible on the galaxy map. This origin is 2x available, so it can be used with a friend in multiplayer!
Utopia
An origin made for "one system challenges"/tall empires. You start with more special and powerful planets.
Arcane
You start on a giant "Arcane"-ringworld that has 8 habitable segments with unique districts. Designed for tall empires/"one system challenge".
Gray Tempest
This is a playable Gray Tempest empire that is based on nanites. You have access to a new trait, ships, technologies, planets, buildings, structures, crisis path & jobs.
Why ask?
Maybe terraforming normal ringworlds to spectralic ringworlds can be an option =)