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Soul gates only open when X amount of guardians have been defeated where X can be anything from 1 to 9. Secondly, bosses now give an item after being defeated. Some, not all, of these items are required to make progress.
In the case with Fafnir, defeating him will unlock a soul gate in Yggdrasil.
Yes, Fafnir locks about everything aside from some exploration past Nidhogg that doesn't give anything that'll early unlock things. Weapon upgrades are much appreciated because leather whip can be a struggle in hard mode