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from guilding land manual:
You lose levels based on monster exclusivity. Eg. Fighting Coral Pukei levels Coral massively but potentially at a cost to all other zones, because that's a Coral-exclusive monster. Hunting Nargacuga would level Coral and Forest, with a heavy bias towards the zone that Narga was actually calling home.
Hunting multi-zone roamer/invader monsters, like Banbaro, Ebony Odogaron, Fulgur Anja, Velkhana, R. Nergi, or S. Deviljho will never de-level any regions for this reason.
As for how high you can level zones, when you start off the cap is 4, as your MR increases you gain Assignments that raise your MR cap as well as the cap for GL zones, eventually capping at 7.
If you want to hunt a monster without changing your region levels, or are happy with your current region levels and don't want them to change for a while, you can Fix the region levels at the Handler/Quest Board.
https://steamcommunity.com/sharedfiles/filedetails/?id=2004179434
Might dunk Wildspire and raise Coral. That would give me the best spread of elders in The Guiding Lands.