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Anyway, once you get enough knowledge points, you advance knowledge level 1 rank and the next row of skills becomes available. Sometimes individual skills will also stay locked behind certain quests being completed. It will either tell you what quest to complete or it will just say "progress the story" which means just keep doing main central quest ring until you get the quest you need.
If you look on the left side of the Mastery screen it says what your current knowledge rank is and how many knowledge points you still have to collect to get to the next rank.