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The largest punishment for missing the beat is that enemies still move. So if you stand aside a red dragon and missed the beat, you may be toast. Even if enemies are not in direct vincinity, you lose initiative where multiple one can start to trap you. Although this only happens in later levels where more enemies spawn.
Also, yea. Without the heart beat, it would probably be to easy. I mean you can test it out with Bard or No Beat mode. It is basically a pure puzzle game at that point. Which is nice but would probably be less interesting.
Consider it like the difference between Chess and Speedchess. Everyone can find the right move if you have enough time (and know the rules). Doing the same under pressure though is a different story. It is still the same puzzle but you have to proceed the information fast enough.
It is also a lot more stylish with the heartbeat and the music. Certainly would not have been as successful as it is like this. Especially considering that the whole story is about the music and rhythm.
- Coin multiplier loss
- Enemies move and you don't
- Obsidian equipment bonus is lost
- Certain items are destroyed
- If it's in a boss fight, you lose the bonus chest
Like it or not, gold coins matter not only for the score, but to properly buy items from the store and hidden ones. Once you start doing All Zone runs you will start to see how important gold is.
Now in the case of certain characters, like Aria or Coda, you die. Pretty sure missing a beat is a punishment. Now add that certain characters have locked mechanics, such as not being able to equip different weapons and you will start to see how everything stacks.
The motivation to play all characters (not in a row) for an All Zones run should be fun or... achievement hunting due to how absurdly hard most of them are.