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As for a comprehensive list, try fextralife.
Stepping Foot in Liurnia
• Skips Roderika into Roundtable Hold (misses out on Golden Seed)
Picking up the Fingerslayer Blade
• Preceptor Seluvis dies (misses out on various Puppets + Magic Scorpion Talisman)
• Pidia, Carian Servant dies
Stepping Foot on Altus Plateau
• Skips encounter with Patches in Liurnia
• Skips encounter with Rya the Scout in Liurnia
— • Skips
• Skips encounter with Bloody Finger Hunter Yura at Raya Lucaria (misses out on Raptor of the Mists Ash of War)
• Initiates the Radahn Festival at Redmane Castle
— • Skips the Alexander encounter at Gael Tunnel
— • Skips pre-Festival dialogue from Castellan Jerren
— • Locks you out of Leonine Misbegotten & Crucible Knight boss until Radahn Festival ends
Stepping Foot on Mt. Gelmir
— Skips the Alexander encounter on the cliff just south of Carian Study Hall
Stepping Foot in Volcano Manor
• Skips Diallos Hoslow questline (if not met in Academy Gate Town; misses out on Diallos's Mask + Hoslow's Petal Whip if not killed)
• Skips Patches encounter in Mt. Gelmir near Gelmir Hero's Grave
• Skips Rya the Scout encounter at Grand Lift of Dectus (if previously met)
— • Locks out Blackguard Big Boggart's shop (misses out on Boiled Prawn and Boiled Crab)
Kill Rykard, Lord of Blasphemy
• Locks out Volcano Manor assassination missions
— • Locks out Old Knight Istvan mission (misses out on Scaled set + Magma Shot)
— • Locks out Rilegh the Idle mission (misses out on Crepus's Vial + Serpentbone Blade)
— • Locks out Juno Hoslow, Knight of Blood mission if you haven't taken the mission yet (misses out on Hoslow Set + Hoslow's Petal Whip + Taker's Cameo)
— • Locks out Vargram the Raging Wolf & Errant Sorcerer Wilhelm mission (misses out on Raging Wolf set + Gelmir's Fury)
— • Locks out Great Horned Tragoth mission (misses out on Bull-Goat set + Magma Whip Candlestick)
• Locks out Rya's questline (misses out on Daedicar's Woe)
Using the Forge of the Giants
• Locks out Brother Corhyn & Goldmask's questline (if Regression riddle isn't solved; misses out on Corhyn's Robe + Goldmask's set + optional ending)
• Causes Melina not to appear in Frenzy ending (if Frenzied Flame Proscription was used after rather than before)
Kill Maliketh, the Black Blade
• Leyndell Capital becomes Leyndell, Capital of Ash
— • Misses out on Bolt of Gransax (needed for achievement)
— • Locks out farmability of Duelist's Set, Duelist's Greataxe, Duelist's Hammer, and Guardian Garb (Full Bloom)
— • Many, MANY other item misses (too many to list)
Other
• Collecting both pieces of Haligtree Secret Medallion — Latenna dies (misses out on Ancient Somber Smithing Stone)
• Joining Ranni's group — Skips Blaidd ahead (misses out on Finger Snap gesture from Kalè + Somber Smithing Stone from Blaidd after defeating Bloodhound Knight Darriwil)
• Assassinate Juno Hoslow, Knight of Blood — Skips Rya-related dialogue from Tanith
Omensmirk Mask
Deathbed Dress
Lionel's armor set
Duelist armor set
Alberich's armor set
Raging Wolf armor set
Imp Head (Corpse)
-
Battle Hammer
Duelist Greataxe
Black Bow
Bolt of Gransax
Cane Sword
Coded Sword
Gargoyle's Halberd
Hammer
-
Gravel Stone Seal
-
Barrier of Gold incantation
-
Thunderbolt (AoW)
-
Sanctified Whetblade
Cracked Pot
Perfume Bottle
Perfume Bottle (2)
"Flightless Bird" Painting
Two Fingers Prayerbook
-
By My Sword gesture
There may absolutely be more, but this was everything I could find upon researching pre-Capital of Ash. This was one of my biggest worries in terms of missables, so if anyone has any to add, feel free to share.
On my playthrough I last saw Alexander at the Radahn festival. I don't know if I talked to him right after the fight, but I know he fought with me.
I was looking up his quest via the wiki and checked ALL locations I might have skipped his encouter before and after the festival. He's nowhere to be found.
So the only times I saw him was were I helped him out of the hole in Limgrave and at the festival. Never in Liurnia, Gael Tunnel, Mt. Gelmir or Farum Azula.
There was no spawn before the fire giant.
Is it a possibility that his quest is locked after you finished the game/beat the Elden Beast?
Where else would he be? I never got the jar helmet nor his shard...
- Killing Maliketh > Changes Leyndell, several unique items are lost.
- Killing Boc > Gives you the "maidenless wretch" status debuff. You monster.