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Thunder and spirit are similar but have one difference - spirit only affect movement speed, thunder makes enemies' animations play faster and thus some of their states update faster - meaning faster attacks for some enemies.
The only reason they are "skulls" is a reference to another game that partially inspired this mod, where there are different run modifiers you can add that are all difficulty skulls. They are also unrelated to skulls and are just damage mods, and figuring that i wanted to make lots of items fast I didn't want to spend a lot of time spriting 20+ new items - so I just went with that theme and did some quick color swaps that are somewhat alright as far as sprites go.