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Unnecessarily long answer:
When you start your journey in Last Epoch you only have access to the base class passive tree. As you progress through the main story you will eventually reach the End of Time map location, which is technically the start of Chapter 4. The End of Time is reached after defeating The Omen of Silence boss that lies within the Sanctum Bastille, of the Ruined Era.
Once you have reached the End of Time you speak to The Forgotten Knight NPC who will open a portal to the Realm of the Forgotten. At the end of a series of walkways and after fending off your other selves you get to a pedestal surrounded by 3 unique versions of yourself. These are your Mastery class options for your character, with but one path to take, and the choice cannot be undone.
Your Mastery choice grants you a unique Mastery Bonus, a skill unique to that Mastery, and full access to that Mastery’s tree. Investing enough passive points into this Mastery tree will unlock a further skill or skills that are unique to only that Mastery. While your Mastery choice locks you to that Mastery, it does not stop you from putting passive points into the other Mastery trees. However, you can only put enough passive points in the other Mastery trees to unlock half the tree.