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Only two AFAIK. Floating Passage and Anti-Air Deathblow was it I think.
Edit: And weren't there also a couple of missable skills you get from killing optional minibosses or bosses? Couldn't say for sure right now, but that might count too.
I think you might be right, though I don't remember which ones. I know that the Ninjutsus are all earned not learned but I dunno outside of that.
However, I just checked his Steam profile and he has all achievements, except the "All skills" achievement (And the achievement for all achievements of course) so I don't think he missed anything apart from maybe the Vendors.
Yes.
The Sakura Bull, O'Rin of the Water, and the second Chained Ogre are all skippable minibosses which give skills. (There are also four "required" bosses/minibosses that give skills which in some cases are possible to skip, like the Blazing Bull).
Additionally, finishing the final bosses of both Shura and non-Shura routes give a skill each, so you need to do both.