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I don't think any of the final bosses drop them.
He’s under the watchtower room Isshin stayed in. Where the 3 nightjar usually are.
The route guide says he and the fire Juzou reskin in the ablazed outskirts are optional, but both drop prayer beads.
Are there any beads that are completely missable even with the box at the shrine other than those in the optional memories at the estate?
You could rather easily tell if it's incomplete or not by counting the number listed.
I have killed Mizuo (that guy was harder that most bosses) and fire Jouzou.
I have the one in the building where you meet Isshin.
i somehow missed Long Arm Centipede Sen-Un in Senpou Temple, I must have walked right by the building without seeing that you could get in via the roof.
Now I can finally go get my revenge on Genichiro.