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Typically, most people do okay when they have less than 500-ish animals in enclosures. The ones in the tanks don't slow you down much. You may have to fine-tune your own average figure.
A big drain on the game is the guest-pathing, I've had a lot of success in reducing lag by installing very wide pathways and placing plazas/squares/piazzas so the guests don't bump into each other as much. when they do, both parties have to recalculate their paths and often end up bumping into each other again etc etc.
Now bear in mind that habitat space alone isn't a factor in lagging. I have a full map in the arctic circle containing just 6 or 7 habitats spread around with a bunch of stores and many guests and staff around the map. The game is having to do a lot at once.
You will need to find your limitation that lets you play the way you want to. Be it guest count, animal count, or how expansive your zoo is. I had hoped that building a spacious zoo would help distribute the guests so that things wouldn't be so bad, but alas, it didn't work as expected.
I think you hit a good point with the biome - weather effects like rain and snow are computationally expensive for large areas.
I'm trying to get every animal in my current Zoo and yes, it's lagging. But considering all above, that's not really surprising.
This Zoo contains:
- 1205 animals (welness 100%)
- 96 species
- 287 staff (satisfaction 99%) (playing in Dutch, don't know what the correct term is)
- 10.119 guests (satisfaction 94%)
- 106 facilities guests
- 51 facilities staff
- 3 rides
Riverboat 5
Gondola 9
4x4 15
Wondering if I'm gonna make it!
Intel(R) Core(TM) i7-9750H CPU @ 2.60GHz
16,0 GB DDR4 RAM
NVIDIA GeForce GTX 1660 Ti
Settings: High.
So, nothing to fancy. Almost 5 years old.
I'm on a mission to try and fit all species in a zoo, currently got about 1400 and while it is lumpy, it is still working. All the enclosures are as plain as they can be while hitting complete happiness for the animals, so it isn't a visual feast, but it has rides to zip people from one end of the map to the other.