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The "Growl-Clucks" are ambient Puma sou... NOISE that plays A LOT when a puma is in the area.
I wish they would skip the Puma ambient noise and just stick with the much less often heard Puma "call" sound.
They are extremely wary animals. If they know you are there, they hold back and aren't as skittish as lynx on Medveg Taiga. If you rush in after them, they run super fast and may trample you but I dont think they are aggressively out to kill you off like a lion when provoked. Others have said it happened that way. I remain unconvinced.
Maybe someone can explain better their experience hunting them. They basically came across as less skittish oversized lynx to me, rather than a smaller lion in terms of attitude.