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I'd say it's a toss between Red deer, fox and coyote. These are by far the most skittish animals in the game from my experience. That said, each animal type has different ratings ranging from 0 to 9 (you can see this when tagging them with the binoculars). This affects how good they are at hearing and spotting you which means even the easiest animal with a rating of 9 will detect you from very far and be extremely hard to catch.
It seems foxes, coyotes and red deers have a higher base skittishness. They also give more money from what I've seen, perhaps it's linked.