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In order to increase lily rank, characters must fight on the front side, rear side does not count, Let me bring Rom and Neptune back, Neptune and Rom must be on the front row and won a certain fights to gain lily level, but to gain lily benefits, Neptune must be on front line and Rom at the back or vice veras.
You can have a look in the Party > coupling section, each character has unique lily ability on the top right table with the three abilties and the lily level requirement. If those abilities are grey, you see that the lily between those two have not met the lily level.
Although, I feel like there might be a bug in some skills because while I was trying to do "win battle without healing" thingy, The challenge always reset when Neptune heal herself even if the lily level with the rear character was 1, which supposed to be 10.
Not sure if this would help, but these are all I know.
1. Just winning fights (all front character gains lily rank between them)
2. Helping other characters, using buffs or items on them.
3. Using lily couple skills (back) or formation skills (front).
The benefits can be obtained when two characters with enough lily rank are coupled, 1 front and 1 rear. Basic benefit are the support, it can be offensive (STR and INT up) or defensive (VIT and MEN up). The other benefits are some passives the rear character give to front character and vice versa, but they must got enough lily rank to be active. For example, IF can increase EXP gained, or Compa reduces the SP amount required for skills.