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Anyway, you covered it pretty good, the only thing I have to add, or to ask, is what happened to the kid of the guy with the sword and shield?
The bosses names are listed as the achievements for defeating each of them.
From what we can gather from the Mothership computer face thing, it made it clear that:
Everyone on earth hates you.
You will still destroy the earth you are saving if you go back.
You are denying the sustenance of whatever beings they are, you are choosing to protect earth over your supposed home. You are called out on this.
In the story, it seems implied that the main character (lets call him The End) simply fell into earth, after he caused much destruction he was locked up and contained. You notice how the Mothership face challenges you when you bet her, questioning the overall point of your actions as no one will protect them from ''the next one''. At this point, The End crash lands back into earth, and the rabbit man walks off with his child.
It sounds to me like the ''save the world'' end is a constant process of locking The End back up and him breaking out and defeating the next wave of agressors trying to assimilate earth, the old man who can time travel even makes a comment about seeing you in the next cycle.
This question is very philosophical, but The Bunny also said something that makes you think at the end: "so now you know. You must think I'm insane, or maybe you udnerstand me. I think you've changed, and now it's all about what you choose to do" or something in these lines.
It's almost as if Rider (that is the name of our character), according to the bunny, doesn't have to destroy the earth. And maybe rider can continue living somehow.
Also, the idea is (in my opinion) that rider learns the emotional world of earth, which doesn't really seem to exist in his own world. Maybe he values it, and when you choose the save the world ending, you basically are saying that rider thinks that the human world is more valued than his own cold world.
Also, why would you destroy their world just by being in it? And why was Rabbit locked up also in the cell?
Now to me each boss represents kinda of a conscious for our protaganist, Rider or The Stranger
The Chain: The first boss we fought represents Hatred. He is cold and held no remorse to Rider (due to Rider being the destroyer of world. This is similar to what Rider was made to be, a cold-blooded murderer.
The Strap: The second boss represents Insanity. She, similar to Rider, destroyed everything in her path and was later put in her own jail, becoming crazy and used to become a Jailer to kill Rider. Rider was created by the The Star to be used to destroy worlds for survival
The Line: The third boss was to me Perspective. As he was the master of time and experienced the future and past he saw the why's and how's to each action, to Rider's motives. This plays a role in the Final Battle with The Star if you decide to save the World. The Star wants to destroy Worlds to insure survival for itself and its creations while the Jailers are protecting it for it's their home and lives.
The Scale: The fourth boss represented Corruption. He speaks on how you corrupted him, turning him into a mutated disfigured being and now wants to take revenge on you. It is similar to the Final Showdown with The Star because The Star who created Rider also corrupted him by making him believe your purpose is to destroy and that's all you are.
The Hand: The fifth boss represented Determination. In the opening intro when we first face wtih The Hand, we see he has a son and is willing to give his life to protect both the future of The World and his son as well. It gives willpower to keep on fighting, a reason to why we fight. Rider, however is fighting for simple survival.
The Song: The sixth boss represented Peace. When we first meet her she is reasoning Rider to stop and stay here, giving him her faith that Rider can change into someone better than a murderer. We are then given an option to stay or to fight her. Staying with Song reveals that she does care about him and with that ends the game, congratualting with us the Peaceful End. If you decide to fight Song, before he killed her, he first hesistated before finishing her, which to me thinks he wanted Peace afterall.
The Burst: The seventh boss represented Winning. In her boss fight and by The Voice (The bunny man) description to her, she thinks everything is just a game and that she must wins everytime. Rider must win every boss because we are playing the game and in order to survive, we must kill the Jailers
The Edge: The eighth and my favorite boss represented Purpose. When The Voice is giving you a brief description of The Edge he reveals that he has been training to fight for you because he believes it is his fate to fight you, his purpose to stay alive and train to witness you. Rider's purpose to destroy because he was told to and that is all he ever did.
The Beat: The ninth and final Jailer boss represented Remorse. After all the Jailers you killed just to survive, during her fight she questions you to why did you do this, you don't seem like a monster. During her death scene she asks for you to Hold her hand, that she wants you to feel someones life you taken away by holding them until the end.
The Voice: Despite the fact he is not a boss fight he is a Jailer which represents well Voice. The Voice is your bunny rabbit friend who assist you in your journey by giving you a reason to why you have to fight and who the Jailers are. He gave you reasoning to keep fighting but to think about if it's worth destroying the world, that you are his only chance to save his World.
I won't talk about The Star because it created you and you decide if you wanna kill that hand head thing or call out a full invasion plan.
The Rider was sent as a frontrunner by The Star to gauge the planet, its resources, its defenses. The Rider himself is a walking wasteland, as you witnessed after finally getting free, he withers the land just by being there. At some point, The Hand led an army against The Rider and won, capturing him then containing him inside a multi-layered prison floating outside the planet, guarded by the best warriors the planet has to offer.
We don't know how long everyone's been stuck inside that prison, but somewhere along the way, The Voice may have been "betrayed" by The Song. Maybe the design was secretly altered while building it, maybe it was sabotage, I dunno, but he mentioned how she "even locked ME up", implying that The Song has something to do with why he can't leave his own creation. Gradually he grows to miss his own daughter and regret his own creation, until desperation overcame him and he sought to use the very prisoner they're supposed to be containing to break both of them out.
The Chain is the cruelest among them, so he was assigned to be the first Jailer, constantly torturing and killing the immortal Rider to keep him weak. He also acts as an alarm, signalling the other Jailers when The Rider finally escapes his cell, as implied by the little purple waves that radiate from each stage when he slammed the ground and the screen panned to the prison's entirety, to show how many levels there are. The Strap's origins aren't clear, but she's been driven completely insane and is clearly powerful, so she was used as another Jailer. Both are simple obstacles in your way, other than feeling a little pity for The Strap you kill them and move on.
The Line is where it gets interesting. He's not a sadist, he's not crazy. For a wise old man to volunteer himself as Jailer, he has to have a pretty good reason. He also foresaw things about The Rider, and apparently The Rider's arrival on the planet interfered with his clairvoyence. On top of that, remember his dying words. "Look to the sky, there you'll find your answer." Or something like that. The Voice also mentions how The Line told them to notice "the dot in the sun", but "nobody listened", except The Voice maybe. What's up in the sky? The Rider's mothership. He also apparently only spoke of these things after the prison was built, as The Voice complained about how he knew, but didn't do anything to stop it, and didn't warn him about how he'd be trapped as well. Why? Maybe The Line gave in to his own sense of fate, maybe he knew if anyone was going to stop The Star, it'd be The Rider himself.
The Scale was mutated by some kind of freak accident and went crazy. He accuses The Rider for being the one who did that to him. Was he just insane? You might think that when you first meet him, but knowing the truth, he's probably not lying. How was he corrupted? Freak accident caused by The Rider? Just by going near The Rider? Maybe. But then why aren't the other Jailers and the environment inside the prison being corrupted? Maybe The Voice created a suppression field against The Rider's withering energy? Who knows.
The Song attempts to reason with The Rider. She offers a small world of nature uncorruptable by The Rider for him to roam free in, even offers herself, in exchange for The Rider staying. Even when The Rider fights her, she heavily expresses her reluctance. Is it mind games like The Voice claims? Maybe, until you defeat her and she simply accepts her fate, no deperate struggle, no final words of defiance. Her death also shows the fact that The Rider isn't just an empty husk or a bloodthirsty killer, he has a conscience of his own, he's capable of hesitation, of recognizing compassion and mercy. For the player, you start to realize that maybe there's something else going on here, her offer of peaceful co-existence was genuine, she actually cares about the Rider. If she's here as a Jailer, then there has to be a very good reason why the Rider's locked up, right? Also The Voice's comments about her are probably mostly out of spite, since she's the one who got him stuck here and away from his daughter.
The Hand further cements the theory that The Rider is in fact the "bad guy". You as the player probably sympathize with a father fighting to protect his son, but maybe The Rider did too. Maybe he recognized the honor in The Hand, the nobility in fighting to protect his loved ones, and in relation, the Jailers fighting to protect their people, the strength it gives them.
The Burst is more like The Chain, not really someone sympathetic, just powerful. The Edge however, the strongest of the Jailers, trained for who knows how long solely for the purpose of facing The Rider. Is he fighting for a sympathetic cause? Not exactly. But maybe The Rider could grow to respect him all the same, as a true warrior, and as an example of the potential of this planet's people, what they can push themselves to become with enough determination, their value and how they earned their right to exist.
And finally, The Beat. Compared to everyone else, an amateur, a pushover, even the Voice knows it and put her behind The Edge, the true strongest, on purpose, plus protecting her with a metric truckload of automated defenses AND a super kill-beam for a desperation attack. If The Edge was the strongest, then she is the weakest... but she's still here, putting her life on the line, fighting The Rider to protect her people against impossible odds. She talks of what the Rider has done, how he withers the land, how he's a monster who'll destroy the world, and when she's finally killed, she pleads to The Rider with her dying breath to hold her hand, maybe so she doesn't have to feel like she's dying alone, maybe to show her a sign that The Rider has a semblance of compassion in him, that there's hope that him getting out won't destroy the world.
The Voice congratulates The Rider. He knows what'll happen next thanks to The Line, he knows The Rider will return to his mothership and potentially bring about a world-ending invasion, it was the price a desperate father was willing to pay to see his daughter again. But he also says that The Rider might not be the same person as when he first arrived. How he hesitated when killing The Song, it brought him hope; maybe some of the Jailers got through to The Rider, changed him, convinced him not to go through with his mission.
The Rider leaves his prison and returns to the planet's surface. The player sees why The Rider was imprisoned, quarantined, that everything The Beat said you were was true. But maybe she's not entirely right anymore? The Rider returns to The Star, and makes a choice. Whose world is worth more? His own cold and emotionless world who'd welcome him as a conqueror? Or the world below with its colorful people, the world who the Jailers sacrificed themselves to protect, the world who'd still hate him and see him as a danger, never knowing he saved them?
Personally it was The Beat who convinced me to rebel the most, though The Voice helped a lot too. There's really no objectively right or wrong choice though, whether to save your home world and destroy a world that fears you, trapped you, tormented you, or to destroy your home world and save a free world and its innocent lives. It's all up to perspective.