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It determines spell casts and weapon arts.
It can be increased by leveling up Focus.
Focus also boosts Attunement.
Also MINI SPOILER!!
Seeing Andre is back as a smith he has an option to assign estus which means if you have high healing estus amounts you can then take it out of estus and put it in your Ash estus which refills the mana bar
Usually Estus chugging is frowned upon in PvP, but in DSIII invaders will be able to chug as well as hosts and summons, so PvP will become a bit more similar to Bloodborne's in that regard.