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He was the one that advised Prince Lothric to stay away from fire. He always wanted Age of Dark, never to prolong Age of Fire.
He is prbably somewhere in Anor Londo, with Hollows, Darkwraiths and rest of his fun bunch.
He is also big fan of Aldrich, which would explain how he become Devour of Gods, tieing up loose ends from the first game.
So at somepoint in time the serpents made contact with the royal family, particulalry influencing the Heavenly Daughter Gertrude, leading to her imprisonment for following the heretical serpents.