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Initially, once you have leveled all heroes to 100(or 125 if they have another upgrade) you want to go back to Treebeast and level in steps of 25 as close as you can to 1,000 then drop down to the hero below, Ivan, and level them to 25 levels lower than Treebeast drop to the one below and level to 25 levels below Ivan and carry on the pattern. Once that is set up, as gold becomes available level Treebeast by 25 levels and do the same down the list of heroes then repeat from the top.
Once you can regularly reach zone 250 before ascending, the process changes. At this point you want to move all Gilds onto the first five full damage heroes, Treebeast, Ivan, Brittany, Samurai and Seer. Use the 2 HS degild option to move them. Once this is done similarly to before you start leveling Treebeast to 1,000, but this time you only level on these five heroes rather than the whole list. Don't stop at 1,000 and note that unused heroes between these five count for a step down of 25 levels in the pattern.
Once you start reaching zone 500 before ascending, you will want to move all Gilds onto Samurai and only level him after the base 100 levels in each hero. You will end up moving the Gilds on from Samurai to Atlas eventually, but that is generally into the mid-game so can be left for here.
This tends to translate into
The first 3, who have 450%, Masked samurai, who has 450%, and Forest sage, who has 450%
The first 3 are earliest available, will reach 1000 first, and cheapest. But. As a result. of the 5, will be the weakest once they're all the same level+gild.
Masked Samurai is a Middle ground. All the skill dps, Available early, stronger, and will reach 1000 earlier than most. But not the strongest, Still quickly obtainable and cheaper than most though. By the time you're getting close to affording Dread knight, Masked Samurai can be level 1150 (x25's) (could have more, probably, but not really worth)
That's *a lot*
Forest sage will be even stronger, has 450% again, but, is going to be the most expensive of the five. Still going to be obtained fairly early each ascend, making the gilds worth coming back to each time, but its expensive pouring into him.
That's just Early on, though, Likely.
http://jsfiddle.net/tvvvbhfg/embedded/result/