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There are different bosses and storylines around the DLC's. The vanilla map has 4 bosses as of now.
The giant glowing columns allow you to upload dinos. You can then download these dinos on different servers in the same cluster.
Also ascending by doing bosses.
You start on the island, then Scorched earth, Abberition and Extinction last.
The Centre and Rag are community maps and non canon.
About the Glowing collum, the large ones are obelisks. They are used to switch inbetween servers (uploading chars and dinos).
The smaller versions are supply drop and contain some loot, level restricted bound to their colour.
Its a sandbox, you can decide your own goal like ascending, building a base, taming stuff, waging war (well, if you run private its at best a skirmish between you and your firends).
You could call the bosses "bad guy" but doesn't fit well.
You only can "escape" to other maps.