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As far as i know it's actually a feature that you're able to plant them inside Caves or maybe even Buildings?? (I'm not sure, i have not tried this out). They can grow on fungal gravel and don't need light to survive.
The moss however should grow on any fertile tile in a radius of 8 blocks. Maybe there aren't any fertile tiles near the tree? Try building fungal gravel around the tree if you haven't already and see if it changes anything. Altough i think the moss is spawned upon planting the tree..
Yes they can be planted inside of buildings
On OP's question, have you waited a few days? the moss should start growing back as far as I know. You do need that fungal gravel to accomodate for it.
The roof hole isn't currently necessary. Other trees cannot be placed under roofs, gauranlen trees can, under both thick and thin roofs-- as the OP mentioned it is possible this is a bug, but if it is a feature then gauranlen trees are specially special trees :p
I agree it is sub-optimal to use inside trees for connection due to the building interference. For ideologies with tree connection meme though they get a huge mood boost just for seeing a gauranlen tree (and a penalty for not seeing trees) so a tree can be left untended purely for mood reasons if you have the seeds to spare. works best for dining+production+barracks mega rooms.
Yes there is fungal gravel around the 3 trees ,there is no construction carried trees