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Fun fact, it is canon to the games lore that every single one of those (normal) Vipers you fight is a female of their species. You only ever encounter a single male in the entire game.
second why would it be a joke? look at the bloody thing, its literally just a snake with arms and hands. thats not exactly a 'human feature' every other alien creature we encounter has that too.
Humans: Bipedal, have arms with hands that have opposable thumbs and are warm blooded.
Snakes: None of the above.
Vipers: have both human and snake characteristics. Hence why Tygan says they have human characteristics. Because they do.
He says "without" and "purely reptilian" So they do not have human DNA, and you must have misheard him.
That line from Tygan is simple a joke about how all the snake enemies in this game (other than the Viper king) have boobs.
Please tell me about the last time you have ever seen a snake with pair of boobies?
I'll wait
The "in game" explanation is generally accepted to be that they are a pair of honking great venom sacks. Just ones that are located on the chest area under an armored corset.
Hence the consistent rumors about this games rule 34. Because the internet doesn't need an excuse, but gods help us all when it finds one.
This is the whole thing "although i would expect to be more growing accustomed to seeing the clear intermingling of both human and alien genetic material, i must admit this reptilian creature with such distinctive human features is surprisingly unsettling, invasion-era reports include refrences to a species with some reptilian qualities, but nothing as readily apparent as this..."
But how could that be unsettling?
Also funny.
Of all the creatures in this game they could have tried to make look feminine, why is the snake monster on that list?
If you go looking in the Vipers wiki page you can find concept art clearly showing that this result was, not, an accident. The devs of this game knew what they wrought.
I mean, it's obviously not the actual real answer - the devs wanted to make sexy+scary and someone obviously had a thing for Lamia's.
But when you find the viper king et all, Tygan and Vahlen's notes kinda suggest that the species that make up Vipers had already been MASSIVELY genetically fiddled with before Xcom 1 and certainly Xcom2.
The viper king seems to be the only male any human's seen of this species. We know that Torque's info says she was born in an egg clutch in (Alaska I think?) - so I'm beginning to wonder if the Elders agreed with Tygan when they first met Vipers and, once dominated their race, ensured viable eggs were only females to control their expansion and numbers. They also, probably, provided by them via genetic mutation and something akin to the gene template in the foundry, but stored off Earth and accessed via the portal/ sterile viper eggs that are injected with DNA to begin the development process.
We also know that it seems the Elders also have issues controlling DNA mutation and expression, even with their powers, as some beings seem to have flaws that should have been able to be removed but cannot be, possibly as they have to use human DNA as the glue to hold it all together and that forces odd expressions of traits? One of which, as Vipers are female snake aliens, means that they have boobs. And, yes, considering how far they can spit their venom? It could VERY well be that the human DNA moved the venom sacks to the front... due to a weird expression mechanic of the female genes.
https://www.pfizer.com/news/articles/how_genetically_related_are_we_to_bananas