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In game - the brain (rather than the head) is really small but if you can hit your targte and kill it - no disadvantages at all.
But is actually scowled upon by the game designers.
The game designers, In their wisdom, have made the skulls of most animals kevlar plated to further discourage players from taking that route.
As Alf pointed out, head movements are many and unpredictable but even while they are going on the body remains reasonably static. This is also true in real life hunting situations.
In real life, being hit in the head would drop most animals cold, not nessecarily dead , even if the brain is not actually penetrated. But if it happens in game the animal will run a considerable distance.
Also, if you shoot an animal in the head in real life, the trophy mount would most likely be spoiled, possibly to the point of being unusable.
So while their are no hard and fast rules about head shooting it is not the optimal choice in game.
I thought that on some of the animals you got a minus on the score/ccs for headshots ?
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I think you mean a decrease in the Trophy Integrity, bought about by an extended Wound Time which happens any time an animal is not instantly killed.
Trophy Integrity can also be reduced if you kill an animal with an overpowered weapon from too close a distance.
For example a red deer shot with a 270 at 50m will have a higer Trophy Integrity than a red deer shot at 50m with a 45.70 even if both have a zero wound time.
In some special comps wound time must be zero and trophy integrity 100% to make the animal qualify for an entry.
http://thehunter.wikia.com/wiki/Harvest_Value
CSS is related to the highest possible score that an animal can acheive within its species.
IE a top of the leader board Reindeer with a optimal rack will have a higher CSS than a spike of the same species..
So CCS can be used to establish the true trophy value ( not score ) of a Fox against an Elk or any other animals.
http://thehunter.wikia.com/wiki/CSS
the trophy integrity has no real purpose.
Also on deers, head can be hard to shot because of unpredictable movements.
On boars and bears, I will try to hit the head if the target is facing me, because the heart will be very close to the ground, and not easy to shoot.
If the target is showing me its back, i will probably try to hit neck or brain, though it will be a harder shot.
Even a double lung shot and they run a good a bit off. One in the brain and they drop like a ton of bricks.
With everything else your pretty much better off with the heart/dual lung shots than going for a headshot.
Yes there is a reason not to shoot. Headshots does not make any sence, in 80% of all cases, because the animals' brains are too small, eg. see how big the skull of a moose is and how small compared to that the brain. So mostly you will hit the skull and that means a lot of tracing. Get them lung- or heartshot and it's down, nice and clean in it's tracks! So that would be my suggestion.
I have hade some it´s in the brain, most because of the angel when hunting from a tower.