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In-game, definitely no, except for the 3 Northeast Corridor routes (New York - New Haven, NY - Philadelphia and the North Jersey Coastline) and engines / trains designed for there, usually CSX, but also some Amtrak. Possibly Miami. Some trains, I think the F59s and so possibly the Caltrain as well have alerter, but no signaling (implemented).
The NEC and some of its branches warranted such systems, plus electrification already required extensive cabling, which has spread into many PRR areas as well at some point, e.g. the Altoonaworks page always mentions track signaling being in place around the Horseshoe Curve (but not modeled on the route).
In the Mid and West, as far as I can tell, no. Usually a crew of two rides these trains and signaling is much more foretelling (similar to the British).
Engines tend to run the whole system, and while some railroads - NS, CSX - specifically forbid non-compliant locomotives to lead (the recent UP excursion train was temporarily equipped with necessary in-cab signaling), in the West (from Chicago) there is a fair mix of UP and BNSF, with a bit of KC and the odd CP / CN mixed in, plus the other way around in Canada (and Mexico with Ferromex). Historically there was no agreement about a single system spanning the continent, maybe PTC will rectify that.
The same story is true in Europe except that due to national borders and randomly chosen system parameters, loco & crew change at borders is standard, so it's not exposed.