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But i would say dryad. You have to place your wall anyways to build your other towers and angry nimbus is a good all around dps, who also drenches enemys so you got that annoying onslaught mutation out of the way. And by switching forms, you also have some cc when needed.
Each of those brings a barricade and at least 1-2 DPS options capable of handling ground & air coverage that scale into endgame.
Squire harpoons might also work.
You'd still be looking at a pretty big challenge without using boosts.