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Well if you want (as an example) your darts to hit faster you can go for Orbs to up there attack rate. or if you want them to hit harder on crits then you put in the Marks.
For me personally i dont know which tower would be good to put them in as it feels like you miss a lot off Attack damage this way.
Marks are for those who want Crit instead of raw Defense Power. Mainly only useful on damage-dealing towers, particularly damage-dealing towers that have enough range that they aren't likely to take hits - Skeletal Archers, Sand Vipers, etc.
Depends on what you're using your Direct Command for. If you got fire to set the oil off, then sure - go DP on your archers. If you don't have or don't use Rain of Oil, then use a mark, go crit, and command your Ramsters instead. If you've got Rain of Oil, Mana Capacitor, and Abyssal Mandate, as well as maxed Mana Cap ascension, use a DP/DHP/DSpd Medallion, load it up with Destruction, Vampiric Empowerment, and Rain of Oil, and command both your Archers AND Ramsters.