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You get to play as the side characters only on DLC Contents, not the campaign itself. You can if you use some sort of Mod.
Other than that... Every character (minus Azrael) have a short side mission DLC which helps expanding the lore, the story or filling some gaps from before or after the main game. Batgirl DLC is the longest one and feels more like a prologue to the Arkham series as a whole.
From there you can play as all of the characters in challenge maps.
Hey, if you are planning to buy Arkham Knight it would be best to get the entire series (including Origins). Otherwise you'll be lost in the story and the rest of the series is amazing. For example: Arkham Asylum has the best story IMO.
But as mentioned above, definitely play them all. Including Origins.
I guess you could say there's a predator style boss (I'm talking about Two-Face), but it's kinda pushing it.
But come on, stop spoiling the game for yourself lol.
If you like Batman, you'll probably like the series as a whole. Bah, except maybe Origins' own spin-off ("Blackgate"), which isn't that bad but not the cup of tea of many.
To your second question, yes, they are, in challenge maps. They also have their own mini stories, I believe, with Batgirl having her own campaign as DLC.
Originally in challenges and short plot stories (less then 1 hour each). Currently it's possible to play them in main campaign as well, if you will install specific mods. Take a look here:
https://www.youtube.com/watch?v=Kr6aGv4m-xU
https://www.youtube.com/watch?v=hKa0VhVDllk
Arkham Knight has the best combat and stealth gameplay, and the batmobile is there.
But it's not well balanced, to much forced batmobile gameplay, and no boss.
The story isn't great either.
City is the best in terms of balance, combat and stealth is great, boss are cool, story isn't stellar but is entertaining.
Origins is my personnal favorite because it has the best story and Christmas setting, but the balance isn't great either: too many fights and not enough stealth.
But at least it has boss unlike Arkham Knight.
City is the best in the serie objectively.
The bossfights kinda suck for the most part but to be fair outside of Mr. Freeze and Origins Deathstroke all the bossfights in the arkham games suck so it's basically business as usual. I would say this is the first and only Batman game that fully sells the batman experience.