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I haven't played too much either - just got to the main town finally so i'm hoping they explain it a bit better once i'm further in.
The paths seem to loosely be tied to elemental damage types.
Inherited skills are carried from previous nodes, but not all weapons have the same number of skill slots so you will definitely lose skills along the way.
I can't remember what the other skill type is, maybe innate, but that is only active when on the current node and can't be Inherited by other nodes.