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Arcanas are extra gimmicky power-ups that you earn for reaching high levels with each of the normal characters. If you turn them on after getting a few of them unlocked, you will be able to choose one at the start of the stage, one out of three randomly chosen from those you unlock after you defeat the boss that spawns at 11 minutes, and one more out of three from the boss that spawns at 21 minutes. (In most stages anyway. The times are sometimes different in the short stages). They have a wide variety of effects.