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The Pal Soul can be used for upgrade your pal stats to be higher and stronger.
There have 4 stats and each stats have HP, Attack, Defense, and Work Speed and each one have maximum with 10 levels and total 40 levels for all upgrade (but combat pals tend to have mostly 30 levels with only upgrade HP, ATK, and DEF to max).
+0 meaning your pal not have soul tag along with your pal and more than +0 meaning your pals have already been upgrade by using Pal Souls.
All newest pals by caught or breed start with nothing soul.
You can upgrade your pal stats with Pal Souls by go to Anubis statue. You also can refunded all Pal Souls back to your inventory from your pal and making them to +0 but cost a lot money.
Trying to maximum on each stats (10 levels) needed 10 Small Pal Souls, 6 Medium Pals Souls, and 6 Large Pals Souls.
You can upgrade Pal Souls converted into larger one or vice versa via Crusher (example Small Soul x2 to Medium Soul x1 or Medium Soul x1 to Small Souls x2).
Pal Souls can randomly found by red chests, yellow chests and rare drop by some pals.
This is a very instructive reply, thank you so much.