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Other than that at the end you will be able to collect 13 keys and will need 10 for the ending so you will have 3 spares if you find them all.
While looking for the answer to this same problem, I stumbled upon an 11th world location with another Warden Key in it that wasn't on the list. Go to 1312:7:1504, near the nose of the crashed ship to the NW of the Standard spawn area in the desert (about halfway to the Lost Paradise location) and I found an 11th world key. It's possible there are even more out there besides that one, but none of the guides I've come across so far listed more than 10 world locations.
-921:87:-628 - lava area
1587:51:563 - ship on a cliff
1895:75:268 - near ship near waterfall
1437:75:-757 - circle ruins
1535:22:2220 - northern jungle
716:67:1708 - sand waterfall
-380:28:1493 - fog crater
-303:84:614 - waterfall next to zeolit area
1316:7:1503 - large ship
2375:97:1009 - waterfalls
1257:17:249 - east of ship on a hill, north of aluminum field
I heard there are certanly 12 (and maybe even 13) hidden keys in total and I am sure the full list is somewhere in one of the threads on steam.
It wasn't that hard to find them without guides. First one was the hardest when I didn't know how locator works, as I thought you will have to explore all suspicious places in different corners. But luckily I found the first key on the way to my base with teleporter, when I was ready to give up, right next to the base so I knew now that they are in the open and not hidden in the corners. After that it was easy to find the remaining 9 and then later I found 11th by accident pretty close to my main base.
Basically you check the map, find a biome or a large wreck where you didn't get a key yet and go there and run through the center of it or around the entrance if it has one. Most likely you will find it.