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It's a human settlement that was attacked by demi-humans and leonine misbegotten must've been the demi-human leader or whatever and stole the castle's sword (that sword thing you get after you kill him)
The lore is still there but a lot of it is gotten through context of the levels. For Leonine you can piece together that he was the one leading the rebellion against Castle Morne. Not all bosses have this however, don't ask me to come up with anything for watchdog beyond "hes guarding a catacomb".
So for the lore, when crossing the bridge to the peninsula you should have seen a woman named Irina by the side of the road. She starts a questline all about going to Castle Morne and finding Edgar. If you did this before killing Leonine you can continue the quest with Irina surviving. If you kill Leonine first she'll end up dead, but you can still continue on with the questline to actually start and finish another questline either way.
So there's a questline leading up to this boss and a questline that takes you into Liurnia. That's why I said technically you should visit this place before even finishing Stormveil or very shortly after. Since after finishing Stormveil it drops you out at the grace where you start the 2nd quest. This 2nd quest actually leads you to an ending for the game.
So in short, yes there is lore behind the boss and the questlines tied to it can actually lead you to one of the like 6 possible endings.
Not everything in a game needs to have deep lore in my opinion. I'm perfectly fine with the watchdogs literally just being guardians of catacombs. We have similar things in reality.. like gargoyles that are meant to 'protect' and so on. It's just in Elden Ring those guardians are less of just statues and more of actual magical beings. I also like that they're reused because it shows a consistency between catacombs throughout the land.
More like bad enemy
Oh yeah I wasn't saying you meant it negatively or even positively. Was just adding my opinion to it. Not every single thing in a game needs like deep lore behind it.. I'm fine with Leonine just kinda being a thing that exists and there is a bit of story involved. I'm also fine with Watchdogs just being tomb guardians.. they are cats after all so I want to assume that idea was taken from Egyptian tombs.
The world itself doesn't scale to you like other open world games. Instead the areas are scaled in a way where you can explore most of them from any point, but they will be harder. Like it's recommended you be about level 50 when doing Liurnia, but you could run straight there from the start of the game if you wanted to. As for bosses not being threatening, it really depends on how overleveled you are for the area. If you're like level 50+ in Limgrave then yes it's going to be easy going through everything. What I will agree with is there's very little like middle ground.. the game a lot of times is either too hard or too easy. It's rare to feel like you're on par with everything.
I'm tempted to agree somewhat here as well. A quest log could have been nice because, like you said, Souls games were very linear so it was easy to curate an experience without it. However, on here there's so much to explore and so many NPCs involved that it's very easy to overlook many of them and even flat out forget they exist. There's a lot of NPCs that aren't even useful in like any way.. like Boc is practically useless.. but he has an entire questline. There are others though like Brother Corhyn that are extremely useful, but his questline barely tells you anything. He at one point basically just says "I'm going on a journey" and disappears to another area of the world.. where? Who knows you better just be lucky enough to find him. Without the wiki I wouldn't have found most of these questlines. On top of all that there's a lot of questlines where if you didn't do it or at least do it to a certain point will just completely fail when you kill certain bosses. Then you're just out of luck getting whatever they had to offer.
I get that all of these are optional and not *required* to beat the game, but some are how you get a lot of unlocks.. like Brother Corhyn's questline actually gets you 4 more faith incantations. Sellen's questline gets you like 4 more sorceries as well as like 3 good mage sets. The sorceries from Sellen's questline are pretty much required for any sorcerer as well.
As has been said, once you finish an area the arrows from the grace don't stop. They're permanently there pointing towards where the boss is/was. There's a lore reason why too if you pay attention. Actually, if you pay attention to the story you're lucky you can even see that guidance.
From the Castle Morne storyline, you get that the misbegotten rebelled because they were treated very badly.
Leonines are misbegotten as well. So that dude is probably their leader, due to combat prowess and wielding a sword infused with vengeance.
What gets mentioned several times is that these folks have some insanely penned up rage, probably due to the way they have been treated for a long time and that that rage caused some degeneration.
Apparrently, this has not always been the case, though:
Kenneth Haight wants to repair relations to the demi-humans, for example, since it's the right thing for the Vassals and Rulers of Limgrave to do.
So, at some point, there must have been amiable bonds.
A lot of misbegotten throughout the game are passive and 'pray' to representations of the demi gods.
Godfrey has a lot of lion iconography and his 'spirit anmial' seems to be a lion as well.
There's a Leonine Crusader in an isolated cave in the Snowfields who (successfully) wields the Golden Order Greatsword, so I guess that ties in pretty well.
I'd say their lore is, they were once allies of Godfrey and family, but got left behind when he and the other now tarnished went into exile.
As with so many things, the greater lore is scattershot across the whole game : D