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Florian Cravic (aka Bernie Crane), Niko Bellic, and Darko Brevic were the only 3 survivors of Niko's 15-man squad in the Kosovo war in 1998.
According to Darko (and not denied by Niko), the squad killed civilians of which some were Darko's neighbours, which was his motivation to betray them.
Sometime during the war, the squad was ambushed by enemy forces; Niko escaped. He believed that the squad could not have been ambushed unless someone within the group had sold them out; he travelled back to where the ambush had taken place, and saw a large pit with the squad members dead inside. After picking through the bodies, Niko realised that two others were missing: Florian Cravic and Darko Brevic.
Concluding that one of the two had been the traitor within the squad, Niko vowed to track down and kill the culprit. Unknown to Niko, Florian would flee to Liberty City, and Darko to Bucharest, Romania.
In 2008, Niko arrived in Liberty City, both to visit his cousin Roman, and to track down Florian, whom he had learned was living there. Eventually, Niko tracked Florian down — who had come out as homosexual, and was having a fling with Liberty City's closeted deputy mayor, Bryce Dawkins. He protested his innocence, and Niko was angered to know that he would not get his revenge any time soon — he had no idea where Darko Brevic could be.
Source: GTA Wiki (don't read further, or you will spoil the story)