Snakes are elongated, limbless reptiles of the suborder Serpentes (/sɜ:rˈpɛnti:z/).[2] Like all other squamates, snakes are ectothermic, amniote vertebrates covered in overlapping scales. Many species of snakes have skulls with several more joints than their lizard ancestors, enabling them to swallow prey much larger than their heads (cranial kinesis). To accommodate their narrow bodies, snakes' paired organs (such as kidneys) appear one in front of the other instead of side by side, and most have only one functional lung. Some species retain a pelvic girdle with a pair of vestigial claws on either side of the cloaca. Lizards have independently evolved elongate bodies without limbs or with greatly reduced limbs at least twenty-five times via convergent evolution, leading to many lineages of legless lizards.[3] These resemble snakes, but several common groups of legless lizards have eyelids and external ears, which snakes lack, although this rule is not universal (see Amphisbaenia, Dibamidae, and Pygopodidae).

Living snakes are found on every continent except Antarctica, and on most smaller land masses; exceptions include some large islands, such as Ireland, Iceland, Greenland, and the islands of New Zealand, as well as many small islands of the Atlantic and central Pacific oceans.[4] Additionally, sea snakes are widespread throughout the Indian and Pacific oceans. Around thirty families are currently recognized, comprising about 520 genera and about 3,900 species.[5] They range in size from the tiny, 10.4 cm-long (4.1 in) Barbados threadsnake[6] to the reticulated python of 6.95 meters (22.8 ft) in length.[7] The fossil species Titanoboa cerrejonensis was 12.8 meters (42 ft) long.[8] Snakes are thought to have evolved from either burrowing or aquatic lizards, perhaps during the Jurassic period, with the earliest known fossils dating to between 143 and 167 Ma ago.[9][10] The diversity of modern snakes appeared during the Paleocene epoch (c. 66 to 56 Ma ago, after the Cretaceous–Paleogene extinction event). The oldest preserved descriptions of snakes can be found in the Brooklyn Papyrus.

Most species of snake are nonvenomous and those that have venom use it primarily to kill and subdue prey rather than for self-defense. Some possess venom that is potent enough to cause painful injury or death to humans. Nonvenomous snakes either swallow prey alive or kill by constriction.

Etymology
The English word snake comes from Old English snaca, itself from Proto-Germanic *snak-an- (cf. Germanic Schnake 'ring snake', Swedish snok 'grass snake'), from Proto-Indo-European root *(s)nēg-o- 'to crawl to creep', which also gave sneak as well as Sanskrit nāgá 'snake'.[11] The word ousted adder, as adder went on to narrow in meaning, though in Old English næddre was the general word for snake.[12] The other term, serpent, is from French, ultimately from Indo-European *serp- 'to creep',[13] which also gave Ancient Greek ἕρπω (hérpō) 'I crawl' and Sanskrit sarpá ‘snake’.[14]

Taxonomy
See also: List of snake genera
All modern snakes are grouped within the suborder Serpentes in Linnean taxonomy, part of the order Squamata, though their precise placement within squamates remains controversial.[15]

The two infraorders of Serpentes are Alethinophidia and Scolecophidia.[15] This separation is based on morphological characteristics and mitochondrial DNA sequence similarity. Alethinophidia is sometimes split into Henophidia and Caenophidia, with the latter consisting of "colubroid" snakes (colubrids, vipers, elapids, hydrophiids, and atractaspids) and acrochordids, while the other alethinophidian families comprise Henophidia.[16] While not extant today, the Madtsoiidae, a family of giant, primitive, python-like snakes, was around until 50,000 years ago in Australia, represented by genera such as Wonambi.[17]

Recent molecular studies support the monophyly of the clades of modern snakes, scolecophidians, typhlopids + anomalepidids, alethinophidians, core alethinophidians, uropeltids (Cylindrophis, Anomochilus, uropeltines), macrostomatans, booids, boids, pythonids and caenophidians.[18]