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FOUR NEEDLES UNDER PLENTIFUL LEAVES - Needles are painful if stabbed, and the experience is not easily forgotten, while falling leaves symbolize passing time. Four Needles feels conflicted because their peers seem to have buried their memory, forgetting that Four Needles even existed (who they once openly praised), consumed by their own selves.
TWO SPROUTS, TWELVE BRACKETS - Two different sprouts: the Ancients, and the civilization that came before them, who built their home in the deepest parts of the world. Same goal. Two Sprouts believed the latter failed, and that their results would differ from their predecessors, having created twelve Iterators to find the Solution — twelve brackets separating the two.
DROPLETS UPON FIVE LARGE DROPLETS - Water vapor fusing with water droplets, becoming the ever-persistent Rain. Droplets observed intently, wishing to understand the Great Cycle. After failing to fully ascend, they now had only one thing to do: observe and ponder fate.
So, after considering everything that has been said, then comparing them to my own theories, here's what I think the names mean: (credit to everyone for their ideas, as I could not have come up with the following on my own!)
NINETEEN SPADES, ENDLESS REFLECTIONS - Spades are designed to dig. The Ancients dug deep to get to the void fluid, and now Nineteen Spades has to spend eternity reflecting on their decisions.
SIX GRAINS OF GRAVEL, MOUNTAINS ABOUND - Grains are thought to be small, or not very significant. Six Grains felt a "mountain" of pride when their peers found a way to break the endless cycle.
A BELL, EIGHTEEN AMBER BEADS - Bells are often painted as either a soothing sound that invites deep thought, or an alarming one that marks time. A Bell felt that it was "their time" to ascend, but their failure to do so resulted in a long period of time in which they must think deeply about what has passed. They are "trapped in amber" in a sense, like prehistoric insects.
Well, the void fluid is technically a liquid, and other substances without a solid form tend to diffuse throughout a given space. In some religious views, it is thought that when you die, your spirit doesn't take a particular form. So, when the Ancients started to ascend, they started losing their bodies' physical forms, and thus their consciousness got spread out across the cavity in which they contacted the void fluid. A few Ancients in particular were able to mostly retain their physical form, though solely because they had more trouble letting go than the rest of their peers, so they could not even half-ascend like the others who got stuck, and are merely an echo with a lesser level of partial-ascension. They still wouldn't be able to keep the solidity of their body, so the essence of their peers (who are still there, just invisible to slugcat) would still flow through them, essentially being "one" with them.
This could be wrong, but I've been thinking a lot about their names lately, and it almost seemed like the numbers could represent their position in society. Nineteen, for example, is the highest number we've heard the Ancients use. Nineteen Spades stated that they were once the "Count of two living blocks." Meaning, they were in charge of two different cities in the sky (as they've built their cities on big "blocks" of material), and these blocks were "alive" with people who had residence there. In basic words, they were pretty high up in the social ranks.
Or maybe the numbers represent how many Ancients are trapped inside one form, as if there were multiple people in one body. The echoes often use "we" or "us," so it makes sense. And for how difficult and risky the process of ascension was said to be, I doubt there were only six of them in total that couldn't properly ascend, when there were thought to be so many of their kind. But how did they merge to become one?
He sure does act like a jerk if you stay too long in his chamber, though!