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So yes, it doesn't matter which one you use.
Edit: Okay I looked it up and seems both of yours slots are 32x so doesn't matters.
But yes always refer to the motherboard manual as each board will handle how all of this works, differently.
When i look at the PRIME Z370-P M.2 Manual i see the following:
So i see no indication of any restriction concerning PCIe.
On AMD boards however you want to make sure not to use the slot connected to the PCIe x16 slots. So its probably the M.2 slot further away from the CPU and closer to the Chipset, as weird as that may sound.
its fine to use any of the m.2 slots on the mobo
the only ssd that runs hot is the kingston rgb ssd
There is no evidence to that. Quite the opposite with all the motherboard that have only one NVMe M.2 Port which normally is the top one.
if it starts the test rig on fire or you need cooling fans for the ssd it has severe problems