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There are also sometimes unique 'command variants' like the green tunic from TOS and Picard's command jacket in the later seasons of TNG.
In the STO timeline, red is still the command branch color, so therefore ship commanders should be wearing red. But with the Odyssey outfit standardization, they added a command variant that has the white shoulder color.
So if you're trying to be 100% accurate to the STO Starfleet uniforms, the ship commander/ranking officer (doesn't necessarily have to be a captain, but does have to be in the command branch) would wear the command variant, which always has the red color.
There's even a guide on the STO Arc webpage for the exact colors, and even more guides on the wiki for canon/non-canon uniforms.
Of course, I say let their aribitrary uniform code be damned and wear whatever you want, but that's just me.
http://www.arcgames.com/en/games/star-trek-online/news/detail/6005853
a commanding officer indeed has white shoulders and the color red.
Though even the game is inconsistent in this propably due to content spanning five years of development, much of it predating Odyssey uniform release.
As far as actual Star Trek canon goes, it has varied wildly over time and between series/movies.
My characters I dress as I like.
Other than ToS, it is red for command/helm, gold for security/operations/engineering and blue/teal for science/medical.
I base this on the TNG and so forth era of trek. Kirk / Spock trek had the colors reversed for operations and command.
http://static.comicvine.com/uploads/original/3/30334/1135487-trek554.jpg
This is inline with the sto uniform code (commanding officer being always red), the uniform color represent the post one holds, not the "career choice".
In those cases there were one case of TNG yellow and two cases of TNG blue. There are tons of examples of commanders that weren't wearing red.
In an interview that I could not verify, several Star Trek writters were asked why all captains wore red. Their responce was; red is recognized as the color of command and although Captains ultimately have authority of which uniform they choose to authorize on thier vessel or station, and could stay in yellow or blue, it is the expectation that they will change to red.
There are alot of examples where Starfleet uniforms vary from ship to ship, station to station or station to ship. (See DS9 Season 1 Episode 1)
In the modern era "mostly."
Starfleet subdivides its men and women and "others/both" into "branches."
Helm/Navigation/Security is Tactical
Engineering/Operations is Engineering
Science/Medical are "Science"
There are various combinations of badges in those groupings. The badges are "Operations", "Science", "Medical" and "Command."
Command winds up being officers chosen(and realistically NCOs for mundane jobs and the Marines) for particular consideration to advance into higher level operational command.
You have to figure the majority would come from the more tactical minded career fields, one of the more apocryphal "Fanon" stories on Kirk is he started out as security but was told by an insturctor to switch to helm b/c "you have command potential."
Everyone was wearing Red, he wore some funky pastel in ST-TMP b/c "disco" baby.
Edit: Ah, here we go. Jumpsuit too was red.
http://www.therpf.com/gallery/data/655/medium/UniformGroup2.jpg
http://www.arcgames.com/en/games/star-trek-online/news/detail/6005853
I've even gone a little further and when I assigned T'Vrell to be my first officer in the duty officer assignments department head stuff and I changed her into the command colors too. It's nifty to know if you like to blend in and/or roleplay somewhat in the game.
However, if you don't care about such things...just wear whatever you like. But bare in mind you can't change the uniform of random crew when you visit the bridge and interior of your ship.
but what with the Odyssee stuff and all, I gave up and now I generally have indistinct grey/ black (or other subtle colors) for my crews overall.
It really doesn't matter that much at all, unless you are heavy into roleplaying.
Then again, I'm always happy to see "decent" uniform setups as opposed to the usual MMO "as eye cancer inducing as possible" mentality so may MMOers seem to favor.
If it's canon, all the better.
Again, not that it matters much at all, but I'm personally more inclined to interact with people who at least try to look "good" in the sense of trying to look Star Trek. Conversely, I pretty much ignore everyone who thinks this is WoW with maximum glitter or what have you.
I have to admit I laughed hard at that Homer Simpson guy running around on ESD sometimes (looks pretty good and accurate, as far as the character creator allows), but that's all it's good for: A laugh. I can't imagine actually running around like that.
Breaks what little immersion the game has.