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The only known thing is they're usually on slopes and not in super high elevations so keeping to locales of that type might be your best bet.
I'll keep trying that, then. I found a lot on the slopes between the Lost and Found, and town. In fact I found most of them when I was often carrying a skull or antler to a new location to hoard for a collection, so i'd leave map markers to mark where to return since you can only hold one thing at a time. Yet I never found a wolf skull, and it led me to going to the far left of the map and even up to the hunting lodge hoping maybe one spawned around there or in the mountains around it, but so far no luck.
Still, for anyone reading this, keep me posted if you find one on the map, and let me know where yours was. I saw someone find one in town, but I myself had no luck. Still, it was worth a shot checking. Here's hoping there is one somewhere on the map and that the game didn't forget and just generate everything else/too many duplicates, lol.
I suggest checking wolf territories and keeping scent view on, maybe you'll come across it in one of the territories.