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The route split happens before the Mega evolution, and at the point of the route split, you can choose between your highest and second highest Karmas. For example, if you have Moral at 40, Wrathful at 39, and Harmony at 20, you'll be able to choose between the Moral and Wrathful routes.
Now, while Champion and Ultimate depend entirely on your current Karma, Mega depends entirely on your route. In the example I gave, you have the option to start the Wrathful route even though your highest Karma is Moral. This will automatically lock your Mega to the Wrathful Mega, even if at the point of evolution your highest Karma is still Moral.