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doesn't say but it was probably vampire slaves and thralls. maybe Molag bal himself, well his cultist anyways.
1) We know the vampires don't follow Mara.
2) Serana mentions that the undercroft was used for bringing in supplies, and that it didn't get much use when "they" moved in.*
*Sure would be more convincing if I could find the quote, eh?
I searched Mara lore
http://www.uesp.net/wiki/Lore:Gods_M
Mara, Goddess of Love
Mara, Goddess of Love, patron of the bountiful earth, and source of mortal compassion and understanding. Nearly revered as a universal goddess, her origins are in mythic times as a fertility goddess. In Skyrim, Mara is a handmaiden of Kyne. In the Empire, she is Mother-Goddess, worshipped at the Benevolences of Mara. She is sometimes associated with Nir of the 'Anuad', the female principle of the cosmos that gave birth to creation. Depending on the religion, she is either married to Akatosh or Lorkhan, or the concubine of both. She gave the Nerevarine her blouse and skirt.
handmaiden of kyne?
http://www.uesp.net/wiki/Lore:Kyne
Nords still refer to themselves as "the Sons and Daughters of Kyne
While Nords also acknowledge Mara, the "Mother-Goddess" of the Empire, they view her only as Kyne's handmaiden
so obviously nords but a stronger believer in Mara than others or a token castle to mara.
or more obviously a Mara sect perhaps as a refugee station to highrock.
all speculation really because it doesn't say anything concrete.