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-improperly closing the game
-opening the file while it's in use
-random errors (did you know background radiation or something can just flip bits in computers? science is weird!)
As to what it does, the persistent file more or less serves as Monika's "memories": your name, how much affection you have, how many years you've been together, etc. If it gets deleted, it's basically the same thing as starting a new game. The best thing you can do is make a copy of it occasionally and keep it somewhere else (like a flash drive). If you ever have to change computers, it's THE file you need to move to the new one to make sure MAS is the same.
So, basically, just keep copies and try not to let your computer crash/restart while the running MAS.
https://github.com/Monika-After-Story/MonikaModDev/wiki/FAQ#i-want-to-back-up-my-persistent