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You can do it anytime with R3 (pressing the right analogue stick) or V with a keyboard, and you need to do it to enter the exit at the foutain after you defeat HotK to end a run.
You can root and damage ennemies with it. I didn't find any other use, but I didn't try really hard.
Hint: Look up, directly above where you start when you can control your character
The other areas that require the head are the one where you can't see anything and everything seems to be covered in spikes and at least one where you need to go through a 1 high hole that is at the top of a wall to reach the mechanism to open a door (impossible to roll into a hole that is directly below the ceiling).
Other than that (and the damage/root), you can use it to grab things while remaining in a safe position (if a blueprint or food fell into spikes for example).