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Trees and Bushes will displace Grass
Grass will displace Lichen
Lichen will grow on 0% soil quality
Everything else has a minimum and wont even root until that minimum is reached.
If the soil quality drops below the minimum things die.
Grass, Trees and Shrubs all self spread (all at different rates). They also improve neighboring hexes soil quality(again at different rates).
Bushes and trees can help quite a bit though because they improve the quality a lot more. So if they take root near the border it will improve the quality quite a bit, which allows grass to spread more easily, which then makes conditions for the bush/tree to spread.