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Perhaps we could have the I or the V group be a unique high priority group that is more detached from the queen (IE ants specifically in that pheromone group ALL bunch up together and stay bunched up together, not unlike the scouting or rapid response groups we see certain ants use in nature). This way you can have a little more control over your ants without tipping the whole balance cart over or having to implement a bunch of new systems.
I'll be responding to this, because there is another strategy that works. There is another way to get ahead. I have beaten insane on 2.1 Rising Tide. Don't read anymore if you want to figure out the strategy on your own.
First, Use your Queen! Figure out how to exploit the Queen's massive health bar. The Queen's health makes her a great shield against rover beetles. Let your Queen take the damage from the rover beetles. You need to risk this.
Second, get those rover beetles! The ant colony needs underground food before night falls. So, be very aggressive, and dig to the weakest rover beetles, with only the two ants. Then, let the rover beetles march to your queen. Your queen will take most of the damage.
Third, make Formica Rufas early! Formica Rufas will be your damage dealers. The queen and the workers will take the damage from the rover beetles. But, your Formica Rufas should win this fight. The ant colony will suffer losses, but the sacrifice is not in vain.
The dead rover beetles will replenish the ant colony. But, now... the seeds are free from rover beetles, and that should be enough food to expand your ant colony. Rinse and repeat until nighttime.
When the hermit crabs arrive, the ant colony should be able to overwhelm the hermit crabs. And, if you do this right, on the first night, a meat wall of hermit crabs will feed the ant colony. Good luck. Hope you can beat it!
Once the second day starts I start taking down the beach tiger beetles closest to my base and clearing the food they guard, and then begin clearing below the first ramp if I still have time before the tide comes in. Once the nighttime creatures spawn I usually try to grab a crab or two between waves spawning, at this point the only serious threats are the spiders if they get lucky/unlucky with target selection or accidentally training a bunch of stuff to my nest by not paying attention.
At this point I can usually just snowball to victory with little danger.
The first time I beat insane with the challenge was on Rising Tide. I don't mind replacing ants if I can get the food back. I lose ants, but more food will be coming into the nest then the food lost. My primary goal is to expand the nest quickly as possible and then at night time turn the hermit crabs into a buffet. In the day time I hunt the tiger beetles. For larve, it's better to keep part of the map under control, clear the areas near the nest, but dont bother clearing every place of larve on the map.
It would seem no priority targeting is planned. Your best bet atm is to just "get ahead," so that you still win even given the worst possible scenario. I'd try for as many as 20 spitters for the third wave of the first night.
You might be able to similarly solve the pheromone problem with just having more workers, or alternatively more combatants so that the delays upon fighting a handful of enemies aren't nearly as severe. Personally, I can't get anything done on the second or third days without first killing as many of the enemies in the areas I want to go to as possible.
There are also some tactics for mitigating randomness and inconsistency. You can spend your entire first day and night without going out to the dangerous areas of the beach. The starting food plus what's not guarded right outside of the nest to the east and south is enough to permit you to harvest food from the underground alone (up to four chambers if you're quick). Also, make sure you're getting that additional chamber the first night, right after you beat the first wave of crabs. While you seem to favor saving those chambers for later, I would advise against this, as having as strong an early-game as possible will roll over into the mid and late-game, allowing you to collect much more food during the day.