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You are right, even I think they are nothing like any of our animals, and even if they could be somewhere in our world im 100% sure that they will be their own category because they are so diverse that is imposible to just be a variation of any other creature, but also is really fun to think about this, thinking about how they work basing on known animals is always a good exercise ^ ^
And they're definitely quadrupeds in my opinion. Again think squirrel or raccoon, they walk on four legs, but can stand on two while using their front legs for something else. The way they slide around all upright in-game looks so awkward to me, that's not how that spine and leg structure work :P
But of course, the more specialized the nichelings gets, the less squirrel or raccoon like they are. A water specialized nicheling would probably be more like an otter and a bearyena hybrid would be more canine like. Stuff like that. But the average "base" nicheling is, to me, definitely squirrel or raccoon like.
As far as I am concerned, nichelings are nichelings. Just like a raccoon dog is neither a raccoon, nor a dog, but a separate kind of animal even though it looks similar to other things.