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Having the gene in both slots and/or on both parents does increase your chances of getting it. It's generally a good idea to have it on both parents for things like snouts and body types, because having it only on one means it might not show due to being recessive.
So if both parents have the basic snout in both gene slots and you're trying to mutate in the Big Snout on both parents for the extra smelling power, you could end up with the baby having it in the recessive gene slot, or if you're lucky, they'll have it in both and have the gene active.
with recessive blind/ shortsighted eyes.
So as Fortuna said it is basically a third gene that the baby can inherit :)
You can selectively breed your animals for Big Nose (the gene which gives superior smelling capibilities), or you can invite wanderers (random animals) with the gene into your tribe.
By selectively breeding the gene, hopefully you use animals with more copies of the gene, such as with big nose, where it is dominant to a few others, there is a 3 in 4 chance the baby will also have big nose if both parents are heterozygous. 1 in 2 if only one parent is heterozygous.
Summary:
This game mostly uses simple Mendelian domaint recessive genetics.
Mutations are fixed once you have set them, and the mutated gene selected has a 50% chance of appearing in the offspring's genetics..