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http://metrovideogame.wikia.com/wiki/Dark_Ones
Anyway, thanks for the infomation, guys.
Difference between the book and game: The Dark Ones aren't evil. I don't think Metro 2033 really ever tried showing this, but in the book, before they launch the missiles and destroy the hive, the Dark Ones scream out to Artyom about how they only wanted to live alongside the humans, not to harm them. To make a better world. Artyom takes his final breath of tainted, surface air, before returning to the Metro, possibly to die. I don't know, Metro 2034 isn't in English yet.
I read that part on the wiki. I think 4A intentionally made the Dark Ones more evil than they are in the book, so you can feel more of a sense of accomplishment and less of a sense of guilt after finishing the game, even if you've reached the bad ending where you blow up the looks-cruel-but-actually-peaceful mutants into pieces. This change made the good ending a bit unreasonable though, because as you said, other than some of Khan's lines, there's absolutely no way to even speculate they are not actually evil. But since 4A managed to cover the 'good' side of the Dark Ones in Metro: Last Light, I'd say I'm satisfied with how they are depicted in the game afterall.
I'm starting to feel I should check out the book someday. It's only getting more intresting. Let's hope my English is good enough. :)