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I'm going to take a stab at one which might fit with your play style.
As you get an epic EM, or any improved quality of machine, just replace the first one in any line of them. Place it first so it gets first pick of the ingredients. Then, since it's producing more than the one which was there, deconstruct from the end of the line until the throughput of the line is back to 'normal'. Then the deconstructed stuff, machines, inserters, etc., can be returned to get their own round of upcycling. Or, just used to make the next part of the factory you're working on. You won't have to redesign any of the stuff built, just chop off the end.
Once you have reduced all the lines in one factory in this way, you could redesign that entire factory to use less space. If you're going to take quality all the way to legendary, however, you could keep things at the new level and just repeat the process as even better machines become available. Either way you're putting the new and improved things to work much sooner than waiting until you have enough to redesign the whole thing and replace it. Just make each little piece a little bit better in tiny bites.
So really the only advantage is a smaller footprint where this especially shines is when designing a tight blueprint using beacons and machines. So take any product for arguments sake circuits we can look at how many wires we need per second. If you cannot reach the speed required with speed modules a higher quality machine will of course produce more wires per second thanks to the faster crafting speed.
Fulgora in particular has a bit more practical application of quality since you cant just place block after block of factory, space is a premium their, even after you get foundations unlocked those are fairly expensive and take time to manufacture at a volume that makes an impact.
This is the main benefit actually. It allows you to indiscriminately stuff buildings like EM-plants that have high power draw, full of productivity modules. And you'll need less speed modules to compensate for the speed loss, allowing you to compensate the power consumption increase via efficiency beacons instead.
This is one of the more convenient ways to take advantage of having a handful of buildings that are a little more quality than the rest.