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I only played once with powers but the important thing is that when you reach mission 5 you need to have 13 runes to spend to unlock and fully upgrade possession for the achievement.
I had a save backup before spending the runes for possession as I never needed that power once the achievement was done.
Yes.. your coins and upgrades are reset. If you collect all the blueprints, you can apply those upgrades at any time you want as long as you have the coins to afford it. But yes, you have to collect loot on every playthrough.